iForest - Biogeosciences and Forestry


Scale dependency of the effects of landscape structure and stand age on species richness and aboveground biomass of tropical dry forests

Gabriela Reyes-Palomeque (1), Juan Manuel Dupuy (1), Carlos A Portillo-Quintero (2), José Luis Andrade (1), Fernando J Tun-Dzul (1), José Luis Hernández-Stefanoni (1)   

iForest - Biogeosciences and Forestry, Volume 16, Issue 4, Pages 234-242 (2023)
doi: https://doi.org/10.3832/ifor4239-016
Published: Aug 23, 2023 - Copyright © 2023 SISEF

Research Articles

The structure and diversity of plant communities respond to changes in landscape structure and vary with spatial scale, stand age and plant size. Therefore, it is important to identify the scale (grain size and extent) at which secondary forest attributes of large and/or small plants and landscape structure are more closely associated. We performed multi-scale analyses in which different grain sizes and extents were assessed to determine the most appropriate spatial scale for assessing the association of large/small tree aboveground biomass and species richness with successional age and landscape structure using regression analysis. AGB and species richness were more strongly associated with landscape structure when large grain sizes (500 m2) were used, with R2 values between 0.31 and 0.43. Variation in AGB and species richness was explained primarily by successional age and landscape structure, respectively. At large extents, successional age was related to the AGB of large trees (R2 = 0.43); at intermediate extents, landscape structure was related to the species richness of large trees (R2 = 0.31). The approach and results of this study may facilitate the identification of appropriate areas and scales for the maintenance or restoration of tree diversity, carbon storage, and the provision of ecosystem services in tropical dry forests.


Scale Effect, Grain, Spatial Extent, Multi-scale Analysis, Secondary Succession, Landscape Metrics.

Authors’ address

Gabriela Reyes-Palomeque 0000-0002-0525-948x
Juan Manuel Dupuy 0000-0001-7491-6837
José Luis Andrade 0000-0002-4991-5020
Fernando J Tun-Dzul 0009-0003-2749-6843
José Luis Hernández-Stefanoni 0000-0002-9559-7131
Centro de Investigación Científica de Yucatán A.C., Unidad de Recursos Naturales, Calle 43 # 130, Colonia Chuburná de Hidalgo, 97205 Mérida, Yucatán (Mexico)
Carlos A Portillo-Quintero 0000-0002-2222-1467
Geospatial Technologies Laboratory, Department of Natural Resources Management, Texas Tech University, Box 42125, Lubbock, TX 79409-2125 (USA)

Corresponding author

José Luis Hernández-Stefanoni


Reyes-Palomeque G, Dupuy JM, Portillo-Quintero CA, Andrade JL, Tun-Dzul FJ, Hernández-Stefanoni JL (2023). Scale dependency of the effects of landscape structure and stand age on species richness and aboveground biomass of tropical dry forests. iForest 16: 234-242. - doi: 10.3832/ifor4239-016

Academic Editor

Paola Mairota

Paper history

Received: Oct 12, 2022
Accepted: Jun 26, 2023

First online: Aug 23, 2023
Publication Date: Aug 31, 2023
Publication Time: 1.93 months

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